Human Performancemedium#916
The 'leans' occur when:
A
the aircraft is in straight and level flight
B
the semicircular canals fail to detect a gradual roll, then the correction feels like a roll in the opposite direction
CorrectC
the pilot is dehydrated
D
turbulence is severe
Explanation
The leans occur when a gradual undetected roll is suddenly corrected, and the vestibular system interprets the correction as a roll the other way.
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